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按:1992年出版的《中非植物志》Flore d'Afrique Centrale兰科卷第2分册中的美冠兰属Eulophia检索表有重要参考价值。但原文是用法文撰写的,不便使用。现用通义千问翻译该检索表,以资参考。
1. Stems with 1–2 well-developed leaves at flowering time, distinctly petiolate, often transversely blotched with white, yellow, or brown …………………………………………………………………………..…… 2
1. Stems with more than 2 leaves, sometimes underdeveloped at flowering, sometimes gradually narrowed into a long petiole, never blotched with white, yellow, or brown ……………………………………………..… 5
2. Petals 4.5–8 mm wide; epichile with lateral lobes auriculate in shape, the median lobe making up about two-thirds of the total length of the epichile ………………………………………………… ..3. E. saundersiana
2. Petals up to 4.5 mm wide; epichile different, with the median lobe making up at most half of the total length
……………………………………………………………………………………………………………… 3
3. Spur not thickened at the apex; epichile 8–12.5 mm wide, with the median lobe deeply bilobed…………….
………………………………………………………………………..…………………… 4. E. lubbersiana
3. Spur with a subglobose thickening at the apex; epichile up to 8 mm wide, with the median lobe emarginate.4
4. Labellum slightly convex; spur with a subglobose thickening at the apex; epichile without ornamentation, with lateral lobes obliquely subtriangular, the median lobe making up about one-third of the total length of the epichile …………………………………………………………………………………… 6. E. ugandae
4. Labellum clearly convex; spur narrowed in the middle; epichile ornamented at the base with two smooth and prominent ridges, with rounded lateral lobes, the median lobe making up about half of the total length of the epichile ………………………………………………………………………………………5. E. maculata
5. Sepals at most twice as narrow as the petals, the latter at least 3.5 mm wide ……………………………… 6
5. Sepals either as wide or slightly narrower than the petals, or half as narrow as the petals but then the petals up to 3.5 mm wide ………………………………………………………………………………………..…33
6. Sepals at least 4 mm shorter than the petals ………………………………………………………………… 7
6. Sepals only slightly shorter (up to 2 mm), equal in length or longer than the petals………………………..16
7. Leaves gradually attenuated into a petiole; petals 20–35 mm long; epichile 25–40 mm long and 20–35 mm wide ………………………….……………………………………………………………………………… 8
7. Leaves not attenuated into a petiole; petals up to 20 mm long; epichile up to 20 mm long and 15 mm wide
. ……………………………………………………………………………………………………..………11
8. Epichile obscurely trilobed, ornamented with 4–5 thickened ridges toward the apex …….… 31. E. latilabris
8. Epichile clearly to deeply trilobed, ornamented with 3–4 ridges from the base or towards the center ………9
9. Epichile broadly conical and abruptly narrowed into a short point…………..…...………33. E. bouliawongo
9. Epichile narrowly conical, gradually tapering into a point …………………………………………….……10
10. Sepals oblong to narrowly elliptic, acute, 3.5–5 mm wide; petals 10–15 mm wide; epichile with ridges continuing as papillae on the median lobe …………………………………..……..………30. E. tricristata
10. Sepals obovate, apiculate to cuspidate, 7–8 mm wide; petals 15–25 mm wide; epichile with ridges not continuing as papillae on the median lobe ……………………………………………..…… 32. E. horsfallii
11. Petals 6.5–10 mm long; spur cylindrical; epichile as long as wide, ornamented from the base with a dense bundle of longitudinal, verrucose ridges that become increasingly prominent but disappear suddenly near the apex, with narrow lateral lobes ………………………………………………….……10. E. hirschbergii
11. Petals 10–20 mm long; spur with a conical base; epichile longer than wide, differently ornamented, with rounded lateral lobes ……………………………………………………………………………….……… 12
12. Spur conical, 3–4 mm long …………………………………………………………………………..…… 13
12. Spur with a conical base then subcylindrical, 5–12 mm long …………………………………..………… 14
13. Spur narrowly conical; epichile clearly trilobed, ornamented from the base with prominent longitudinal ridges…………………………………………………………………………………….. 22. E. coutreziana
13. Spur widely conical; epichile slightly trilobed, ornamented from the base with 3–5 entire or slightly crenulate ridges ……………………………………………………………………………… 14. E. speciosa
14. Spur curved to recurved; epichile about 5 mm wide, ornamented with about 5 verrucose ridges almost to the apex ……………………………………………………………………………………… 9. E. rugulosa
14. Spur nearly straight to slightly curved or recurved; epichile 7–12 mm wide, ornamented with 3–7 longitudinal ridges, first crenulate then prominent, narrow and smooth on the median lobe but disappearing before the apex ………………………………………………………………………………………..…… 15
15. Sepals 8–10 mm long; petals 15–20 mm long; labellum slightly and inversely sigmoid; spur nearly straight or slightly curved, 8–12 mm long; epichile 15–18 mm long …….…… 8. E. orthoplectra var. orthoplectra
15. Sepals 5–7 mm long; petals 10–12 mm long; labellum slightly sigmoid; spur slightly recurved, 5–6 mm long; epichile 10–12 mm long ……………………………………… 8. E. orthoplectra var. schweinfurthii
16. Sepals slightly shorter than the petals ………………………………………………………………..…… 17
16. Sepals as long as or slightly longer than the petals ………………………………………………..……… 21
17. Sepals 13–18 mm long; petals 17–20 mm long and 10–12 mm wide; epichile 15–20 mm long, ornamented from the base with 3 longitudinal and lobulate ridges, the median lobe making up about one-third of the total length of the epichile …………………………………………………………...……… 24. E. calantha
17. Sepals up to 8 mm long; petals up to 10 mm long and 8 mm wide; epichile up to 10 mm long, differently ornamented, the median lobe making up from one-half to two-thirds of the total length of the epichile ….18
18. Labellum slightly and inversely sigmoid; spur 4.5–6 mm long ……………………………………………19
18. Labellum different; spur 2–4.5 mm long …………………………………………………………….…… 20
19. Sepals 4–6 mm long and 1–1.5 mm wide; petals 5–7 mm long and 3.5–5 mm wide; epichile 5–7 mm long, ornamented at the base with 2 lateral crenulate protuberances, then with 5 fine ridges disappearing before the apex ……………………………………………………………………………...……… 25. E. arenicola
19. Sepals 6–8 mm long and 3–4 mm wide; petals 7–9 mm long and 7–8 mm wide; epichile 9–10 mm long, ornamented from the base with 5 longitudinal divergent ridges, first strongly crenulate then gradually attenuating and disappearing toward the middle of the median lobe ………………………… 11. E. sylviae
20. Sepals about twice as narrow as the petals; labellum convex; spur subcylindrical, 2.5–4 mm long; epichile with ridges becoming increasingly prominent but disappearing suddenly before the apex ..13. E. clitellifera
20. Sepals 3–4 times as narrow as the petals; labellum concave to slightly sigmoid; spur widely conical, about 2 mm long; epichile with ridges gradually disappearing toward the apex …………….… 12. E. tuberculata
21. Leaves gradually attenuated into a petiole …………………………………………………………...…… 22
21. Leaves not attenuated into a petiole ………………………………………………………………….…… 23
22. Perianth bicolored; sepals greenish and marked with brown; petals dark yellow; labellum dark yellow, clearly convex; epichile 10–15 mm long and 6–10 mm wide, ornamented with a longitudinal smooth ridge, the median lobe clearly convex …………………………………………………………. 7. E. streptopetala
22. Perianth whitish, pink, or purplish; labellum sigmoid; epichile 18–40 mm long and wide, ornamented at the base with 2 short lateral ridges, the median lobe different ………………………………… 35. E. cucullata
23. Epichile ornamented with 2 ridges or 3–5 longitudinal smooth ribs ………………………………..…… 24
23. Epichile with different ornamentation …………………………………………………………………… 26
24. Labellum clearly and inversely sigmoid; epichile ornamented with 2 ridges gradually tapering toward the apex, lateral lobes auriculate in shape ………………………………………………... 26. E. livingstoniana
24. Labellum convex; epichile ornamented with 3–5 ribs and lateral lobes not auriculate ………..………… 25
25. Sepals 10–12 mm long and 2.5–3 mm wide; petals 10–12 mm long; epichile 15–20 mm long, the median lobe making up about two-thirds of its total length; gynostemium 4–6 mm long ……..… 28. E. malangana
25. Sepals 20–35 mm long and 4–6 mm wide; petals 18–25 mm long; epichile 20–25 mm long, the median lobe making up about half of its total length; gynostemium 12–15 mm long ……… ...29. E. flavopurpurea
26. Peduncle with a central scale; epichile ornamented with 3 lines of papillae …………… 66. E. williamsonii
26. Peduncle with 2–6 scales; epichile with different ornamentation ………………………………………… 27
27. Spur subcylindrical, rounded, about 1.5 mm long and 1 mm in diameter; epichile ornamented from the base with 3 longitudinal, verrucose ridges, arched at the base of the median lobe and occupying the central one-third of its length ………………………………………………………..…………… 16. E. fernandeziana
27. Spur sacciform or conical, 3–5 mm long and 3–8 mm in diameter at the base; epichile with different ornamentation ……………………………………………………………………………..……………… 28
28. Epichile ornamented either centrally with 2 short and thick ridges, or from the base with 2 longitudinal ridges continuing into 5–7 lines of fine and dense papillae ………….…………………………………… 29
28. Epichile ornamented from the base with 3–5 longitudinal ridges reaching almost to the apex ……..…… 31
29. Perianth purplish; spur about 8 mm in diameter at the base; epichile obscurely trilobed, ornamented centrally with 2 short and thick ridges, the median lobe making up about one-fourth of the total length of the epichile ……………………….………………… ……………………………………27. E. coeloglossa
29. Perianth of variable color but with yellow-papillose labellum; epichile clearly trilobed, ornamented from the base with 2 longitudinal ridges continuing into 5–7 lines of fine and dense papillae, the median lobe making up about one-half of the total length of the epichile ……………………………………………… 30
30. Spur 3–5 mm long and 4–7 mm wide, wider than long, blunt, and terminating in a short, recurved tip…….
………………………………………………………………..…………… 21. E. caricifolia var. caricifolia
30. Spur 4–5 mm long and about 3 mm wide, as long as or longer than wide, acute …………………………
………………………………………………………………….……21. E. caricifolia var. angulennisepala
31. Leaves linear, 3–6 mm wide; peduncle bearing contiguous scales; sepals acute to long-cuspidate; labellum green and white, striped with violet, with an inferior profile distinctly convex; spur narrowly conical, about 6 mm long and 4 mm in diameter at the base; epichile clearly trilobed; gynostemium 15–18 mm long……..
……………………………………………………………………….……………………… 34. E. laurentii
31. Leaves oblong to narrowly elliptic, 7–50 mm wide; peduncle bearing spaced scales; sepals oblong to spathulate, subobtuse to rounded; labellum generally yellow or sometimes whitish to rosaceous but with dark yellow ridges, sigmoid; spur sacciform to broadly conical, 3–4 mm long and 5–8 mm in diameter at the base; epichile obscurely to slightly trilobed; gynostemium 7–15 mm long ……………………..…… 32
32. Epichile ornamented with 3 prominent ridges at least 1 mm high ……… 20. E. angolensis var. angolensis
32. Epichile ornamented with 4–5 prominent ridges less than 1 mm hig….. 20. E. angolensis var. mweruensis
33. Spur absent or sacciform and indistinct ………………………………………………………..………… 34
33. Spur present and distinct, conical or ± cylindrical ……………………………..………………………… 39
34. Labellum ± articulated; spur sacciform; epichile slightly trilobed, ornamented centrally with 2 smooth lateral ridges and apically with some papillae ……………………………………………… …….37. E. alta
34. Labellum not articulated; spur absent to nearly absent; epichile different ………………….…….……… 35
35. Epichile trifid, with lateral segments oblong and parallel to the median segment …………..…………… 36
35. Epichile entire or trilobed, with lateral lobes rounded to subrounded and perpendicular to the median lobe..
………………………………………………………..…………………………………………………..…37
36. Sepals about 13 mm long; petals about 12 mm long; epichile ornamented from base almost to apex with numerous papillae ………………………………………………………………………… 39. E. schaijesii
36. Sepals 28–50 mm long; petals 20–40 mm long; epichile ornamented only near the apex with numerous papillae ……………………………………………………………………………………… 38. E. walleri
37. Bracts not exceeding the flowers; labellum 6–7 mm wide, with the median lobe ornamented at the base with a zone of numerous filiform papillae disappearing near the apex, making up about one-fourth of the total length of the labellum ………………………………………………………………… 40. E. suzannae
37. Bracts exceeding the flowers; labellum 8–15 mm wide, ornamented at the base with 2 ridges, the median lobe making up about two-thirds of the total length of the labellum …………………………………….. 38
38. Labellum 12–15 mm wide, the median lobe ornamented at the base with 3–7 bundles of verrucose ridges…
……………………………………………………………………………………………… 1. E. eustachya
38. Labellum 8–12 mm wide, the median lobe practically unornamented ……………… 2. E. ruwenzoriensis
39. Spur 1.5–3 cm long; epichile 2–3 cm long ……………………………….……………………………… 40
39. Spur less than 1.5 cm long; epichile generally up to 2 cm long or, if longer, then the spur is no more than 5 mm long …………………………………………………………………………………………………… 41
40. Leaves long-petiolate, appearing at the beginning of the flowering period.36. E. guineensis var. guineensis
40. Leaves subpetiolate and appearing at the end of the flowering period ….. 36. E. guineensis var. purpurata
41. Spur almost straight and horizontal, first filiform then globular at the apex, 4–8 mm long; epichile very short, with truncate, fimbriate apex ornamented with a forward-projecting smooth tongue-like structure…..
……………………………………………………………………………………...……….… 44. E. gracilis
41. Spur and epichile different ……………………..………………………………………………………… 42
42. Sepals both up to 2 mm wide and at least 4 mm longer than the petals …………………..……………… 43
42. Sepals not both up to 2 mm wide and at least 4 mm longer than the petals ……………………………… 50
43. Spur with a cylindrical part at least 5 mm long, never subglobular at the apex …………..……………… 44
43. Spur with a cylindrical part up to 4 mm long, sometimes subglobular at the apex ………………….…… 46
44. Sepals at least twice as long as the petals; labellum convex to slightly and inversely sigmoid; epichile 18–20 mm long, ornamented from the base with 3 longitudinal and lacinate ridges, the median lobe making up nearly two-thirds of the total length of the epichile …………………………………...…… 57. E. paradoxa
44. Sepals less than twice as long as the petals; labellum different; epichile up to 12 mm long, ornamented with 3–5 longitudinal ridges turning into papillae on the median lobe, which makes up about one-third to one-half of the total length of the epichile ……………………………………………….…………………… 45
45. Leaves present during flowering; pedicels not both erect and curved; lateral sepals with auriculate base on the lower side; epichile slightly trilobed …………………….………………………… 63. E. penduliflora
45. Leaves appearing outside the flowering period; pedicels erect and curved; lateral sepals without auriculate base on the lower side; epichile clearly trilobed ………………………………………… 58. E. katangensis
46. Spur subglobular at the apex; epichile ornamented from the base with 3 longitudinal ridges gradually continuing into irregular lamellae on the median lobe, which makes up about half of the total length of the epichile ……………………………………………………………………………………… 59. E. venulosa
46. Spur not subglobular at the apex, at most slightly thickened; epichile with different ornamentation, the median lobe making up less than half of the total length of the epichile ……………………….………… 47
47. Pedicels about 5 mm long; sepals at least 15 mm long; spur subconical; epichile 8–10 mm wide, with the median lobe making up about one-third of its total length ……………………….……… 56. E. longisepala
47. Pedicels 10–20 mm long; sepals no more than 15 mm long; spur subcylindrical; epichile 5–7.5 mm wide, with the median lobe making up nearly half of its total length …………………………………………… 48
48. Peduncle 2–2.5 mm in diameter; sepals 15–25 mm long ……………….…… 50. E. seleensis var. seleensis
48. Peduncle 1–1.5 mm in diameter; sepals 10–15 mm long ………………………………………………… 49
49. Spur almost straight ………………………………………..……………… 50. E. seleensis var. tenuiscapa
49. Spur curved backward ………………………………………...………… 50. E. seleensis var. kisanfuensis
50. Leaves generally coriaceous; spur conical then narrowed and finally ellipsoidal, strongly curved forward, 7–10 mm long; epichile 15–20 mm long, ornamented with 3 longitudinal ridges that are thin at first then lobed, lateral lobes almost as long as the median lobe ………………………….…………… 44. E. petersii
50. Previous characters not associated …………………………………………………………………...…… 51
51. Lateral sepals slightly auriculate and appendaged internally at the base; lip appearing articulated on the auricles of the lateral sepals……………………………………….……………………………..………… 52
51. Lateral sepals neither auriculate nor appendaged at the base; lip not appearing articulated on the lateral sepals………………………………………………………………………………………..……………… 53
52. Sepals 8–10 mm long and 1–1.5 mm wide; petals 1.5–2 mm wide; epichile approximately 8 mm long and 2.5 mm wide, entire, narrowly subelliptic, ornamented from the base with five dentate longitudinal ridges extending from the posterior half to the apex………………………………….…………71. E. callichroma
52. Sepals 10–16 mm long and 2.5–3 mm wide; petals 3–4 mm wide; epichile 12–16 mm long and 8–12 mm wide, distinctly trilobed, ornamented with three longitudinal ridges continuing as papillae nearly to the apex
…………………………………………………………………………………...…… 61. E. tanganyikensis
53. Sepals and petals twisted upon themselves; lip nearly straight; spur horizontal, about 3 mm long; epichile ornamented from the base with three irregularly dentate longitudinal ridges……… 55. E. subsaprophytica
53. Sepals and petals not twisted upon themselves; lip different……………………………………………… 54
54. Leaves shortly attenuated into a petiole, narrowly elliptic to elliptic, 1–3.5 cm wide; spur ellipsoidal, 3–7 mm long and 1–2 mm in diameter; epichile ornamented with a single longitudinal ridge from base to near the apex, subentire or slightly ciliate………………………………………………………… 41. E. euglossa
54. Previous characters not associated and spur never ellipsoidal……………………………..……………… 55
55. Flowers vertical; pedicels erect; lip distinctly and inversely sigmoid; spur saccate at the base, about 5 mm in diameter, prolonged by a straight, cylindrical, notched appendage 2–3 mm long; epichile ornamented from the base with three more or less prominent longitudinal nerves…………...…………60. E. rhodesiaca
55. Previous characters not associated………………………………………………………………………… 56
56. Spur sac-shaped and conical, about 2 mm long and wide; epichile slightly trilobed, ornamented at the base with a fringed ligule extended by four to five smooth ridges reaching the apex, median lobe making up about two-thirds of the total length of the epichile………………………………….……… 42. E. gonychila
56. Previous characters not associated and epichile with different ornamentation…………………………… 57
57. Sepals 15–35 mm long and 2–6 mm wide; petals 15–25 mm long and 4–7 mm wide; spur narrowly conical, 4–5 mm long; epichile 12–20 mm long and 10–15 mm wide, obscurely trilobed, subovate, ornamented from the base with three to five closely spaced longitudinal ridges, these turning into a dense cluster of papillae on the median lobe………………………………………………..……………… 51. E. galeoloides
57. Previous characters not associated………………………………………..……………………………… 58
58. Epichile ornamented on the median lobe with 3–7 closely spaced, verrucose longitudinal ridges……… 59
58. Epichile with different ornamentation…………………………….……………………………………… 61
59. Epichile ornamented with five longitudinal ridges gradually tapering toward the middle of the median lobe, the central ridge often shorter than the lateral ones, lateral lobes not surrounding the gynostemium…….
…………………………………………………………………………………………………15. E. parrilla
59. Epichile ornamented with five to seven longitudinal ridges progressively higher and abruptly disappearing before the apex, lateral lobes surrounding the gynostemium…………………..………………………… 60
60. Leaves present at flowering; flowers pendulous; sepals about 12 mm long; petals nearly 10 mm long; lip convex; spur cylindrical and curved; epichile with the median lobe forming about one-half of its total length……………………………………………………………………………… 18. E. vleminckxiana
60. Leaves absent at flowering; flowers spreading; sepals and petals about 4 mm long; lip slightly sigmoide; spur subclavate, straight; epichile with the median lobe forming about two-thirds of its total length……
…………………………………………………………………………………………… 17. E. letouzeyana
61. Petals elliptic, about 7 mm long and 3.5 mm wide; epichile ornamented with a network of more or less marked veins, slightly papillose at the base but smooth on the broadly obovate, rounded median lobe which forms two-thirds to three-fourths of the total length of the epichile…………………..……. 19. E. lejolyana
61. Previous characters not associated and epichile with median lobe never entirely smooth………………... 62
62. Peduncle bearing at least two scales, either spaced or contiguous…………………………………...…… 63
62. Peduncle bearing generally a single central scale……………………………………………….………… 74
63. Pedicels 20–30 mm long; spur broadly conical at the base and tapering to a point, 4–5 mm long and 3–4 mm in diameter at the base; epichile ornamented with 3–5 crenulated longitudinal ridges... 43. E. crassilata
63. Pedicels up to 10 mm long; spur subcylindrical to cylindrical; epichile with different ornamentation……64
64. Sepals and petals up to 10 mm long; epichile up to 10 mm long…………………………………..……… 65
64. Sepals and petals at least 10 mm long; epichile at least 10 mm long………………………...…………… 68
65. Peduncle 2–4 mm in diameter, bearing 4–8 contiguous scales; bracts subfoliaceous, up to 25 mm long; petals 3–4 mm wide; epichile ornamented from the base with 2–3 longitudinal ridges extending into numerous filaments on the median lobe…………………………………………………… 67. E. kyimbilae
65. Peduncle 1–2 mm in diameter, bearing 2–4 spaced scales; bracts not subfoliaceous, 4–8 mm long; petals 1–2 mm wide; epichile ornamented from the base with 3–5 prominent nerves or ridges continuing as papillae on the median lobe……………………………………………………………………….………………… 66
66. Pedicels about 5 mm long; sepals purplish; lip with almost flat lower profile; spur 5–6 mm long, acute at the apex………………………………………………………………………………… 66. E. saintenoyana
66. Pedicels 1–2 mm long; sepals yellow or greenish-white; lip with slightly concave lower profile; spur 1.5–4 mm long, rounded or with a globular thickening at the apex……………………………………………… 67
67. Herb with leaves; perianth yellow; spur nearly straight……………………..….… 49. E. milnei var. milnei
67. Herb apparently leafless; perianth greenish; spur distinctly curved……...…… 49. E. milnei var. norlindhii
68. Sepals 18–40 mm long; petals 13–35 mm long; epichile 15–30 mm long…………………..…………… 69
68. Sepals up to 15 mm long; petals up to 13 mm long; epichile up to 15 mm long…………….…………… 72
69. Flowers spreading; sepals reddish to brownish; petals whitish and streaked with purple; spur curved forward; epichile greenish and streaked with purple, distinctly trilobed, ornamented from the base with two lateral ridges followed by two rows of papillae, median lobe oblong, forming about one-half of the total length of the epichile…………………………………………………..………………… 23. E. stachyoides
69. Flowers nodding; sepals and petals yellowish, rarely purplish or whitish; spur straight; epichile yellowish, orange, rarely purplish or whitish, obscurely or slightly trilobed, ornamented at the base with two longitudinal ridges, median lobe elliptic or obovate, with scattered papillae usually forming a round, mostly dark purple spot………………………………………………………………………….………………… 70
70. Perianth orange; epichile without colored spot………………………… 47. E. abyssinica var. aurantiaca
70. Perianth not orange, usually yellowish, sometimes whitish or purplish; epichile with a colored spot on the median lobe, usually purple or violet……………………………………………………………………… 71
71. Epichile with median lobe as long or slightly longer than wide………… 47. E. abyssinica var. abyssinica
71. Epichile with median lobe shorter than wide………………………………. 47. E. abyssinica var. euantha
72. Leaves up to 35 mm wide, attenuated into a petiole; inflorescence compact; flowers nodding; perianth essentially yellow but lip with brownish or purplish papillae; spur up to 1.5 mm long; epichile gradually ornamented from base to apex with 2–4 longitudinal ridges continuing as irregular papillae on the obliquely winged lateral lobes……………………………………………………..………………48. E. odontoglossa
72. Leaves up to 5 mm wide and not attenuated into a petiole; inflorescence rather loose; flowers spreading; perianth never yellow; spur at least 2 mm long; epichile different…………………………………...…… 73
73. Upper leaves generally exceeding the inflorescence, which is 2–6-flowered; sepals 4–5 mm wide; spur first conical then subcylindrical, 3–6 mm long; epichile 13–15 mm wide, ornamented with two ridges, median lobe forming about one-third of the total length of the epichile……………………………… 46. E. imbibii
73. Leaves not exceeding the inflorescence, which is 6–10-flowered; sepals 2–2.5 mm wide; spur subcylindrical, 2–3 mm long; epichile about 4 mm wide, ornamented with three ridges, median lobe forming about two-thirds of the total length of the epichile………………………..…… 62. E. trilamellata
74. Sepals 14–20 mm long; epichile 8–12 mm wide; gynostemium about 7 mm long……………….……… 75
74. Sepals up to 13 mm long; epichile up to 5 mm wide; gynostemium up to 4 mm long…………………… 76
75. Inflorescence 3–6-flowered; peduncle about 1.5 mm in diameter; bracts 5–7 mm long; lip flat to slightly and inversely sigmoid; spur nearly straight……………………………………………… 53. E. monotropis
75. Inflorescence 10–25-flowered; peduncle about 4 mm in diameter; bracts up to 20 mm long; lip convex; spur curved……………………………………………………………………………………… 52. E. nyasa
76. Inflorescence fairly compact, 10–20-flowered; flowers pendulous; spur straight, first subcylindrical but with an ellipsoidal thickening at the apex……………………………………….…….…………………… 77
76. Previous characters not associated………………………………………………………………………… 78
77. Sepals yellowish to greenish; spur yellowish to greenish, about 2 mm long………54. E. eylesii var. eylesii
77. Sepals purplish; spur purplish, about 4 mm long…………………………… 54. E. eylesii var. auquieriana
78. Sepals about 12 mm long; petals about 10 mm long; epichile about 12 mm long, ornamented from the base with three longitudinal ridges that are irregularly laciniate…...…………………………… 69. E. acutilabra
78. Sepals up to 10 mm long; petals up to 8 mm long; epichile up to 9 mm long, with different ornamentation...
………………………………………………………………………………………………………………79
79. Leaves about 5 in number, present at flowering; perianth yellowish-green; epichile ornamented from the base with three prominent longitudinal nerves continuing as filiform papillae on the median lobe, which forms about one-half of the total length of the epichile……………………………… 70. E. bouharmontii
79. Leaves generally absent or reduced during flowering period; perianth of different coloration; epichile with different ornamentation, median lobe forming about one-third of the total length of the epichile……… 80
80. Bracts up to 12 mm long and 2.5 mm wide; flowers erect, pedicel about 3 mm long; sepals curved, 9–10 mm long; epichile slightly trilobed, ornamented from the base with a network of prominent veins continuing as small denticulate ridges on the median lobe……………………………… 64. E. chilangensis
80. Bracts about 5 mm long and 1.5 mm wide; flowers spreading, pedicel about 6 mm long; sepals not curved, 7–7.7 mm long; epichile distinctly trilobed, ornamented from the base with three longitudinal ridges continuing as papillose lamellae on the median lobe………………….…………….… 68. E. lambinoniana
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