Regulation of flowering in rice:twoflorigengenes,acomplex gene network,andnaturalvariationRegulation of flowering in rice。这里面的遗传控制图应该是最新的，有新的水稻抽穗期基因出来可以添到图中，如果正在克隆，应该在里面找到相应的位置。

还有篇08年的综述Florigen and the Photoperiodic Control of Flowering in Rice，昨天无意中搜到

Florigen and the Photoperiodic Control

补充水稻抽穗期控制基因

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1 Hd1, a major photoperiod sensitivity quantitative trait locus in rice, is closely related to the Arabidopsis flowering time gene CONSTANS

Hd1,这应该是水稻最早通过图位克隆的方法克隆的一个QTL和抽穗期控制基因

2 Ehd1, a B-type response regulator in rice, confers short-day promotion of flowering and controls FT-like gene expression independently of Hd1

ehd1 这个基因在拟南芥中没有同源基因,是水稻中特有的花期控制途径的关键基因

 Hd1和Ehd1不光和抽穗期有关,还和穗发育相关,见这篇文章

The Flowering-Time Genes Heading date 1 and Early heading date 1 Together
Control Panicle Development in Rice

Panicle formation by flowering-time genes in rice.pdf

既然这样,我当然的大胆假设IPA1也和抽穗期有关了.瞎猜猜,反正猜错了也不用负责,而猜对了以后到能炫耀下.这笔生意稳赚不赔

Regulation of OsSPL14 by OsmiR156 defines ideal plant architecture in rice

Regulation of OsSPL14 by OsmiR156 defines ideal plant architecture in rice.pdf

3 Hd3a, a Rice Ortholog of the Arabidopsis FT Gene, Promotes Transition to Flowering Downstream of Hd1 under Short-Day Conditions

Hd3a 

4 Hd3a protein is a mobile flowering signal in rice

mobile

5 Hd3a and RFT1 are essential for flowering in rice

RFT1

6 Ghd7 Natural variation in Ghd7 is an important regulator of heading date and yield potential in rice

Ghd7

7 Komiya, R., Yokoi, S., and Shimamoto, K. (2009). A gene network for long-day flowering activates RFT1 encoding a mobile flowering signal in rice. Development 136, 3443-3450.

RFT1-

这篇文章确认RFT1是LD条件下的floral activator

8 A pair of floral regulators sets critical day length for Hd3a florigen expression in rice

pair of floral regulators

这对floral regulator分别为:Ehd1(floral promoter)和Ghd7(floral repressor)

9 EL1 Rice early flowering1, a CKI, phosphorylates DELLA protein SLR1 to negatively regulate gibberellin signalling

水稻赤霉素抽穗期调节途径的基因.不过还未见文章明确提及水稻中的该途径

EL1

10 Variations in Hd1 proteins, Hd3a promoters, and Ehd1 expression levels contribute to diversity of
flowering time in cultivated rice

three genes

11 Se13 Complete loss of photoperiodic response in the rice mutant line X61 is caused by deficiency of phytochrome chromophore biosynthesis gene

Complete loss of photoperiodic response in the rice mutant.pdf

12 Ghd8 A Major QTL, Ghd8, Plays Pleiotropic Roles in Regulating Grain Productivity, Plant Height, and Heading Date in Rice.

A Major QTL, Ghd8, Plays Pleiotropic Roles.pdf

 

再来个拟南芥的

Cytokinin promotes flowering of Arabidopsis via transcriptional activation of the FT paralogue TSF

Cytokinin promotes flowering of Arabidopsis via.pdf该研究表明激素cytokinin在花期控制中起作用,另外还有个基因TSF

拟南芥FT-TSF,水稻Hd3a-RFT1

关于细胞分裂素,在水稻产量控制中也有重要作用

Cytokinin oxidase regulates rice grain production

Complete loss of photoperiodic response in the rice mutant.pdf

该基因的克隆也有点意思:Gn1位点有两个基因,Gn1a和Gn1b

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Candidate Gene Association Mapping of Arabidopsis Flowering Time

Candidate Gene Association Mapping of Arabidopsis Flowering Time.pdf这篇文章后面列了n多拟南芥抽穗期控制基因,还有图

进一步补充:

14-3-3 proteins act as intracellular receptors for rice Hd3a florigennature10272.pdf

这篇是分子机理研究的经典之作.以后水稻抽穗期调控基因的文章中估计都会出现FAC.FAC是由Hd3a,14-3-3蛋白,和OsFD1组成的复合体,该复合体结合到OsMADS14的启动子上,启动OsMADS14的表达,启动开花.

The 14-3-3 Protein GF14c Acts as a Negative Regulator of Flowering in Rice by Interacting with the Florigen Hd3a(Plant Cell Physiol. 50(3): 429–438 (2009))

429.pdf

这两篇文章是一个研究团队做的,从PCP到NATURE中间可能有不少故事.

Ehd3:

Ehd3, encoding a plant homeodomain finger-containing
protein, is a critical promoter of rice flowering

j.1365-313X.2011.04517.x.pdf

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FT的新功能:

 

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