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本年初,在PhytoKeys发表了地梢瓜的修订文章。现根据新近看到的一些资料,做些补充与更正。
1、形态描述:在原来审稿中,根据审稿人意见删去了花药、花粉器、种子的过多描述。现补上,并略删减。尤其种子在蒙药里入药,不应过度简化。附上中文描述,下划线标注一些关键特征。
Perennial suffrutescent sometimes lianescent herbs, usually densely shortly pubescent throughout, with white latex, arising from monopodial slightly woody creeping slender brown rhizome up to 3 m × 3 mm, with wiry horizontal offshoots. Stem and branches straight or twining above, green to dark purple; when straight 10–40 cm long, erect or ascending, simple or much divaricately branched from base, with internodes 2–30 mm long; when twining up to 2 m long, little branched, with internodes 4–8 cm long. Leaves opposite or sometimes subopposite, rarely 3- or 4-whorled, with petioles 0.5–10 (–12) mm long or subsessile; blade green, thin, linear, narrowly lanceolate, oblong-lanceolate or occasionally broadly lanceolate, 2–10.5 × 0.1–2 (–2.3) cm, apex shortly acute, obtuse or acuminate, rarely rounded, base slightly oblique, attenuate, cuneate, truncate or rounded, with colleters, margins entire, ciliolate, often revolute; mid-vein elevated abaxially, lateral veins obscure; both surfaces densely pubescent, sometimes glabrate, except the mid-veins on lower surface. Inflorescences alongside the leaf axils to terminal, with 1–10 fragrant flowers, shortly umbel- to raceme-like; peduncles 1–10 (–50) × 0.5–1.5 mm, puberulent; pedicels 1.2–10 × 0.2–0.6 mm, puberulent. Sepals ± half length of corolla, oblong, triangular or lanceolate, 1–2.8 × 0.3–0.8 mm, puberulent, ciliate, apex obtuse, acute or acuminate. Corolla white or greenish-white to yellow, 3–5 × 3–8 mm, usually glabrous, sometimes sparsely puberulent on dorsal surface and inside tube, rarely densely puberulent on both surfaces; tube 0.5–1.5 mm long; lobes 4.3–5.2 × 0.8–1.5 mm, narrowly triangular, lanceolate, oblong or oblong-ovate, apically twisted clockwise, apex ± acute or obtuse, sometimes retuse. Corona of 5 slightly fleshy lobes partly fused at bases, cupular, 0.8–1.8 mm long, shorter or longer than gynostegium; tube shorter than anthers, 0.3–0.7 mm long; lobes oval, triangular-lanceolate to linear, apices erect or incurved, acute, acuminate, obtuse or rounded, 0.5–1.4 mm long, sinuses between theses each with or without a minute triangular tooth. Anthers 5, square, 0.3–0.7 mm long; appendages white, membraneous, erect, ovate-triangular to lanceolate, 0.7–1 × ca. 0.4 mm, as long as gynostegia, 1–2 times the length of anthers, with sparse thick cilia, apex acute or acuminate; pollinaria 5; corpusculum ovoid or ellipsoid, somewhat shorter or half-length of pollinia, 190–200 μm long, 100–150 μm wide, slightly narrowed toward base; caudicles obliquely pendent, 108–150 μm long, thick, attached to apical pollinia, enlarged at the point of attachment to pollinia; pollinia pendulous, oblate or ellipsoid, 228–334 × 119–133 μm. Gynostegia sessile; head conical, bilobed, convex or slightly concave. Follicles paired or single, ovoid-fusiform, ventricose, 4–10 × 0.8–2.5 cm, apex obtuse or attenuated, smooth or colliculate, puberulent; seeds reddish-brown, flat, ovate or subelliptical, 5–10 × 3–5 mm, smooth or sometimes with dark brown granules scattered over one or both surfaces; bottom obtuse-rounded, top truncate, margin wing-like, entire or slightly repand; the concave surface with a slightly elevated and darker line (linear hilum) about 2/3 length of the seed; coma white, 1–2.4 cm long, attached to micropylar end.
多年生灌木状或攀援状草本植物,通常全体密被短柔毛,具白色乳汁;根状茎稍木质,棕色,具坚实的水平侧枝,长达3米×3毫米。茎和枝条直立或盘绕,绿色至深紫色;直立时长10-40厘米,直立或上升,从基部简单或多分散地分枝,节间长2-30毫米;盘绕时长达2米,分枝较少,节间长4-8厘米。叶对生或有时互生,罕3-4轮生,叶柄长0.5-10(-12)毫米或近无柄;叶片绿色,薄,线形、狭长披针形或偶宽披针形,2-10.5 × 0.1-2(-2.3)厘米;顶端短尖、钝或渐尖,稀圆形;基部稍倾斜,渐尖、楔形、渐狭、截形或圆形,具黏液毛;边缘整齐,略有缘毛,常背曲;背面中脉凸起,侧脉模糊;两面密被柔毛,有时除下表面的中脉外脱毛至近无毛。花序沿各叶腋腋生至枝条顶生,1-10朵花,花具淡香味,短伞形至总状花序;花序梗1-10(-50)× 0.5-1.5毫米,被毛;花梗1.2-10 × 0.2-0.6毫米,被毛。萼片约等于花冠的一半长,长方形、三角形或披针形,1-2.8 × 0.3-0.8毫米,被毛,有缘毛,顶端钝、尖或渐尖。花冠白色、淡黄绿至黄色,3-5 × 3-8毫米,通常无毛,有时背面和内面稀疏被毛,稀密被毛;筒部长0.5-1.5毫米;裂片4.3-5.2 × 0.8-1.5毫米,狭三角形、披针形、矩形或矩形椭圆形,顶端顺时针扭曲,顶端±尖或钝,有时截形。副花冠裂片5个,稍肉质,在基部融合,杯状,长0.8-1.8毫米,短于或长于花柱;筒部长0.3-0.7毫米;裂片卵形,三角形披针形或线形,顶端直立或内曲,尖,渐尖,钝或圆形,长0.5-1.4毫米,各裂片间具一三角形小齿或无。花药5个,方形,长0.3-0.7毫米;附属物白色,膜状,直立,卵三角形至披针形,长0.7-1 × 约0.4毫米,与花柱等长,是花药的1-2倍长,顶端尖或渐尖;花粉器5个;着粉腺椭球形或椭球形,稍短于至仅达花粉块的一半长,长190-200微米,宽100-150微米,向基部稍微变窄;着粉腺柄斜垂,长108-150微米,粗,连接到花粉团的顶端并略膨大;花粉块下垂,扁球形或椭球形,长228-334 × 119-133微米。合蕊冠无柄;花柱顶端圆锥形,浅二裂,凸起或稍凹。蓇葖果成对着生或单生,卵球状梭形,膨大,4-10 × 0.8-2.5厘米,顶端钝或变细,表面平滑或具圆形突起,被短毛;种子红褐色,扁平,卵形或近卵形,5-10 × 3-5毫米,光滑或有时在一面或两面具深褐色疣状突起;底部钝圆,顶部截形,边缘翼状,整齐或稍微波状;凹面有一条稍微凸起和较深的线(线状种脐)约为种子长度的2/3;种毛白色,长1-2.4厘米。
2、Cynanchum thesioides 的模式指定:原文指定K.F. Karo 127 (BRNM [15481/36], ex herb. J. Freyn) 为主模式,并认为保存在PR的两份标本K.F. Karo s.n. (BR [PR793598, PR793599]) 很可能不是isotype。尽管这几份标本都是在1871年7月24至8月5日采集的,但无法确定后两份标本是否与BRNM的标本是在同一天采集的。不过,我们最初是将BRNM与BR的标本视为syntypes,因为作者在这一系列的文章开篇提到他看到了3套采集人的标本,我们也正好寻迹找到了作者发表这个新种的3套标本(下图),也因此最初是将BRNM的标本选作lectotype。尽管最终我们是根据编辑的意见进行了修改,但我个人还是持保留意见,觉得编辑对于同一个gathering的定义过于严苛。
图1. 地稍瓜(Vincetoxicucm thesioides Freyn)的模式标本标签。
3、PPBC照片:去年底文章接受时,PPBC中地梢瓜的照片有2000张多一点。一年过去了,又增加了近500张照片,将一些错误鉴定的排除后,现在有2496张照片了。
图2. PPBC中原错误鉴定为地梢瓜的照片(现重新鉴定了)。前3张的重新鉴定,是因为网站系统默认与下面几张变色变前的照片是同一个批的,导致的重复鉴定。
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